If a specific node uses a version of the GETH client where it doesn't check for the users Ether balance when sending Ether, what happens? Would the node just accept a transaction where the user doesn't have enough balance? Would that lead to an underflow with Ether balances?

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No,before being being added to the blockchain other node's/ validator's will also verify whether the transaction is legit.

The validity of that particular transaction is not checked only at originating node

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